A population is a group of individuals of same species that live in a given geographical area, share or compete for similar resources and potentially reproduce.
Ex: All the cormorants in a wetland, rats in an abandoned dwelling, teak-wood trees in a forest tract, bacteria in a culture plate and lotus plants in a pond etc.
Population ecology is an important area of ecology as it links ecology to population genetics and evolution.
Population Attributes
Birth rates: Refer to per capita births.
E.g.: In a pond, there are 20 lotus plants last year and through reproduction 8 new plants are added. Hence, the current population = 28
The birth rate = 8/20 = 0.4 offspring per lotus per year.
Death rates: Refer to per capita deaths.
Ex: 4 individuals in a laboratory population of 40 fruit flies died during a week.
Hence, the death rate = 4/40 = 0.1 individuals per fruit fly per week.
Sex ratio: A population has a sex ratio.
Ex: 60% of the population is females and 40% males.
Age pyramid: It is the structure obtained when the age distribution (% individuals of a given age or age group) is plotted for the population.
For human population, age pyramids generally show age distribution of males and females in a combined diagram.
Population size or population density (N): It is the number of individuals of a species per unit area or volume.
Ex: population density of Siberian cranes at Bharatpur wetlands in any year is <10. It is millions for Chlamydomonas in a pond.
![](https://blogger.googleusercontent.com/img/b/R29vZ2xl/AVvXsEiabNeBAz8XLl3I4ZozZeuVzYh0iQrIilP_vtGqNel02WF_gy0T_54UdikFLyVOdYl58m5DSv3EbPvfWzaQg5NFYAE8greDkeO0qHZjDN17yBOsOpNQK229RrW9_r0rtdfnmD2xe3AydmZ5/w640-h126/pyramid.png)
Population Growth
$\displaystyle \small \bullet$ The size of population is not static. It keeps changing with time, depending upon food availability, predation pressure and adverse weather.
$\displaystyle \small \bullet$ The main factors that determine the population growth are
Natality (B): It is the number of births in a population during a given period.
Mortality (D): It is the number of deaths in a population during a given period.
Immigration (I): It is the number of individuals of the same species that have come into the habitat from elsewhere during a given time period.
Emigration (E): It is the number of individuals of the population who left the habitat and gone elsewhere during a given time period.
$\displaystyle \small \bullet$ Natality and immigration increase the population density and mortality and emigration decrease the population density.
$\displaystyle \small \bullet$ If N is the population density at time t, then its density at time t +1 is
$\displaystyle \small N_{t+1}=N_{t}+[(B+I)-(D+E)]$
$\displaystyle \small \bullet$ Population density increases if B+I is more than D+E. Otherwise it will decrease.
$\displaystyle \small \bullet$ Under normal conditions, births and deaths are important factors influencing population density. Other 2 factors have importance only under special conditions.
$\displaystyle \small \bullet$ E.g. for a new colonizing habitat, immigration may be more significant to population growth than birth rates.
![](https://blogger.googleusercontent.com/img/b/R29vZ2xl/AVvXsEjKA7u-nzROvpeTxK1apeYnyF7E_tBz-XCNq42b6aPPoYdQr9UC_G92fDo18S6_qlB6FKp0tVw690L791x4ONslVRVAxs584MQRd6GmIc6j8J0KRI97ZN4E5jwyiLM0VLT2-p0Vw-FcmXgH/w400-h250/popdensity.png)
Growth model
Growth of population takes place according to availability of food, habit condition and presence of other biotic and abiotic factors.
Types of models
Exponential Growth
$\displaystyle \small \bullet$ In this kinds, growth occurs when food and space is available in sufficient amount.
$\displaystyle \small \bullet$ When resources in the habitat are unlimited, each species has the ability to release fully its innate potential to grow in number.
$\displaystyle \small \bullet$ The population grows in an exponential or geometric fashion.
$\displaystyle \small \bullet$ If in a population of size N, the birth rates as represented as ‘b’ and death rate as‘d’. Then increase and decrease in N during unit period time ‘t’ will be
dN / dt = (b – d) × N
Let (b – d) = r, then
dN / dt = rN
$\displaystyle \small \bullet$ The r in this equation is called ‘intrinsic rate of natural increase’.
$\displaystyle \small \bullet$ The r is an important parameter for assessing impacts of any biotic or abiotic factor on population growth.
r value for the Norway rat = 0.015
r value for the flour beetle = 0.12
r value for human population in India (1981) = 0.0205
The integral form of the exponential growth equation is $\displaystyle \small N_{t}=N_{0}e^{rt}$
where,
$\displaystyle \small N_{t}$ = Population density after time t
$\displaystyle \small N_{0}$ = Population density at time zero
r = intrinsic rate of natural increase
e = the base of natural logarithms (2.71828)
![](https://blogger.googleusercontent.com/img/b/R29vZ2xl/AVvXsEi8bUjcwZFRCG6naL9b0NJextYf10eW_2_JSL9Mm67NbeQ61kpMWuAfBFqoWNjK5YZSulwS2MaBRe8tBWSVB0o_oDxW4wVXGAKLlzyG6gu-5J4xgEuAda3AEjMzsdvE1XMHZ_fm9KjUJUs3/s320/expo.png)
Logistic growth
$\displaystyle \small \bullet$ There is no population in nature having unlimited resources for exponential growth. This leads to competition among individuals for limited resources.
$\displaystyle \small \bullet$ Eventually, the ‘fittest’ individuals survive and reproduce.
$\displaystyle \small \bullet$ In nature, a given habitat has enough resources to support a maximum possible number, beyond which no further growth is possible. It is called carrying capacity (K).
$\displaystyle \small \bullet$ A population with limited resources shows initially a lag phase, phases of acceleration and deceleration and finally an asymptote. This type of population growth is called Verhulst-Pearl Logistic Growth.
It is described by following equation:
$\displaystyle \small dN/dt=rN\left ( \frac{K-N}{K} \right )$
where,
N = Population density at time t
r = Intrinsic rate of natural increase
K = Carrying capacity
$\displaystyle \small \bullet$ Since resources for growth for most animal populations are finite, the logistic growth model is more realistic one.
![](https://blogger.googleusercontent.com/img/b/R29vZ2xl/AVvXsEiLOZ5qgRFuj8PBzEGdv0d_iXIw931LEgMncnVjTyRLc4OD8TU8U07mAsG_sz5mlQm9C7E-KZfbjoYct8PEHNeoKU_aThwnS1fqn7i2xw0yP-DYjKtB6TT0kgb8QMdwZTPb6oZqcOXHjaMD/s320/logistic.png)
Ex: All the cormorants in a wetland, rats in an abandoned dwelling, teak-wood trees in a forest tract, bacteria in a culture plate and lotus plants in a pond etc.
Population ecology is an important area of ecology as it links ecology to population genetics and evolution.
Population Attributes
Birth rates: Refer to per capita births.
E.g.: In a pond, there are 20 lotus plants last year and through reproduction 8 new plants are added. Hence, the current population = 28
The birth rate = 8/20 = 0.4 offspring per lotus per year.
Death rates: Refer to per capita deaths.
Ex: 4 individuals in a laboratory population of 40 fruit flies died during a week.
Hence, the death rate = 4/40 = 0.1 individuals per fruit fly per week.
Sex ratio: A population has a sex ratio.
Ex: 60% of the population is females and 40% males.
Age pyramid: It is the structure obtained when the age distribution (% individuals of a given age or age group) is plotted for the population.
For human population, age pyramids generally show age distribution of males and females in a combined diagram.
Population size or population density (N): It is the number of individuals of a species per unit area or volume.
Ex: population density of Siberian cranes at Bharatpur wetlands in any year is <10. It is millions for Chlamydomonas in a pond.
![](https://blogger.googleusercontent.com/img/b/R29vZ2xl/AVvXsEiabNeBAz8XLl3I4ZozZeuVzYh0iQrIilP_vtGqNel02WF_gy0T_54UdikFLyVOdYl58m5DSv3EbPvfWzaQg5NFYAE8greDkeO0qHZjDN17yBOsOpNQK229RrW9_r0rtdfnmD2xe3AydmZ5/w640-h126/pyramid.png)
Population Growth
$\displaystyle \small \bullet$ The size of population is not static. It keeps changing with time, depending upon food availability, predation pressure and adverse weather.
$\displaystyle \small \bullet$ The main factors that determine the population growth are
Natality (B): It is the number of births in a population during a given period.
Mortality (D): It is the number of deaths in a population during a given period.
Immigration (I): It is the number of individuals of the same species that have come into the habitat from elsewhere during a given time period.
Emigration (E): It is the number of individuals of the population who left the habitat and gone elsewhere during a given time period.
$\displaystyle \small \bullet$ Natality and immigration increase the population density and mortality and emigration decrease the population density.
$\displaystyle \small \bullet$ If N is the population density at time t, then its density at time t +1 is
$\displaystyle \small N_{t+1}=N_{t}+[(B+I)-(D+E)]$
$\displaystyle \small \bullet$ Population density increases if B+I is more than D+E. Otherwise it will decrease.
$\displaystyle \small \bullet$ Under normal conditions, births and deaths are important factors influencing population density. Other 2 factors have importance only under special conditions.
$\displaystyle \small \bullet$ E.g. for a new colonizing habitat, immigration may be more significant to population growth than birth rates.
![](https://blogger.googleusercontent.com/img/b/R29vZ2xl/AVvXsEjKA7u-nzROvpeTxK1apeYnyF7E_tBz-XCNq42b6aPPoYdQr9UC_G92fDo18S6_qlB6FKp0tVw690L791x4ONslVRVAxs584MQRd6GmIc6j8J0KRI97ZN4E5jwyiLM0VLT2-p0Vw-FcmXgH/w400-h250/popdensity.png)
Growth model
Growth of population takes place according to availability of food, habit condition and presence of other biotic and abiotic factors.
Types of models
Exponential Growth
$\displaystyle \small \bullet$ In this kinds, growth occurs when food and space is available in sufficient amount.
$\displaystyle \small \bullet$ When resources in the habitat are unlimited, each species has the ability to release fully its innate potential to grow in number.
$\displaystyle \small \bullet$ The population grows in an exponential or geometric fashion.
$\displaystyle \small \bullet$ If in a population of size N, the birth rates as represented as ‘b’ and death rate as‘d’. Then increase and decrease in N during unit period time ‘t’ will be
dN / dt = (b – d) × N
Let (b – d) = r, then
dN / dt = rN
$\displaystyle \small \bullet$ The r in this equation is called ‘intrinsic rate of natural increase’.
$\displaystyle \small \bullet$ The r is an important parameter for assessing impacts of any biotic or abiotic factor on population growth.
r value for the Norway rat = 0.015
r value for the flour beetle = 0.12
r value for human population in India (1981) = 0.0205
The integral form of the exponential growth equation is $\displaystyle \small N_{t}=N_{0}e^{rt}$
where,
$\displaystyle \small N_{t}$ = Population density after time t
$\displaystyle \small N_{0}$ = Population density at time zero
r = intrinsic rate of natural increase
e = the base of natural logarithms (2.71828)
![](https://blogger.googleusercontent.com/img/b/R29vZ2xl/AVvXsEi8bUjcwZFRCG6naL9b0NJextYf10eW_2_JSL9Mm67NbeQ61kpMWuAfBFqoWNjK5YZSulwS2MaBRe8tBWSVB0o_oDxW4wVXGAKLlzyG6gu-5J4xgEuAda3AEjMzsdvE1XMHZ_fm9KjUJUs3/s320/expo.png)
Logistic growth
$\displaystyle \small \bullet$ There is no population in nature having unlimited resources for exponential growth. This leads to competition among individuals for limited resources.
$\displaystyle \small \bullet$ Eventually, the ‘fittest’ individuals survive and reproduce.
$\displaystyle \small \bullet$ In nature, a given habitat has enough resources to support a maximum possible number, beyond which no further growth is possible. It is called carrying capacity (K).
$\displaystyle \small \bullet$ A population with limited resources shows initially a lag phase, phases of acceleration and deceleration and finally an asymptote. This type of population growth is called Verhulst-Pearl Logistic Growth.
It is described by following equation:
$\displaystyle \small dN/dt=rN\left ( \frac{K-N}{K} \right )$
where,
N = Population density at time t
r = Intrinsic rate of natural increase
K = Carrying capacity
$\displaystyle \small \bullet$ Since resources for growth for most animal populations are finite, the logistic growth model is more realistic one.
![](https://blogger.googleusercontent.com/img/b/R29vZ2xl/AVvXsEiLOZ5qgRFuj8PBzEGdv0d_iXIw931LEgMncnVjTyRLc4OD8TU8U07mAsG_sz5mlQm9C7E-KZfbjoYct8PEHNeoKU_aThwnS1fqn7i2xw0yP-DYjKtB6TT0kgb8QMdwZTPb6oZqcOXHjaMD/s320/logistic.png)
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